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The fishes histories go back thousands of years. As a species it is super interesting... some evolved in hot-spring conditions that would simply cook most other animals. =3
https://en.wikipedia.org/wiki/Red_garra
https://www.youtube.com/watch?v=CLWkojAnyH0
It would be interesting to learn if this occurs with other species of ants. I suppose until now nobody thought to look.
There's a famous paper/framework called "Major evolutionary transitions in individuality" that sketches out a big picture pattern of major evolutionary advances in complexity following a surprisingly consistent pattern: As cooperation and division of labor strongly increase, selection starts working on larger entities. This pattern holds all the way back to the origin of life itself as things moved from self-assembling molecules to compartmentalized populations of molecules, from replicators to chromosomes, from RNA+enzymes to DNA+proteins, from prokaryotic cells to eukaryotic cells, from unicellular to multicellular, from individuals to colonies/superorganisms, and (possibly) onwards to more complex societies
https://www.pnas.org/doi/10.1073/pnas.1421402112
Both are fundamental. You can't survive without cooperating, but you also can't survive if you try to cooperate with the entire biosphere because ultimately there is competition for scarce resources. If you don't assert yourself to claim your share of those, something else will.
That is in no way at odds with what I said.
- picking a good location for a cleaning station requires long-term planning and real strategic judgment;
- deciding which hosts to accept is a complex skill requiring some sort of rudimentary theory of mind + long-term development of social ties;
- like crows, cleaner fish are jerks who constantly try to screw each other over, so there is something of a cognition arms race.
I will add that the wrasse family of cleaner fish use rocks to smash open shellfish (i.e. they are tool-users), and they have very complex group strategies for raising their young. In fact I'm not convinced that wrasse evolved to be cleaner fish at all: they are natural scavengers and scum-suckers, perhaps cleaning stations are a form of cultural technology.
I would be extremely surprised if any of this was true for cone ants. I suspect that is more hard-wired, perhaps a local subspecies stumbled into a genetic fluke, and as you say due to game theory it is a local optimum this population has settled on. If this behavior were common like it is in vertebrates, we probably would have seen it earlier. But who knows? 20 years ago I would have thought "fish have a form of culture" is too ridiculous an idea to consider.
[1] Seriously: https://journals.library.columbia.edu/index.php/cusj/blog/vi... https://www.nature.com/articles/s41598-025-25837-0
OK.
> Cleaner fish are highly intelligent[1]
Yes.
> and it appears that this intelligence is necessary
Manifestly not, at least not in general.
> for their niche:
That is an open question. Just because an unintelligent cleaner fish hasn't evolved doesn't mean it couldn't.